Greenland Shark (Somniosus microcephalus)

Vertebrate Choice Project

1. Taxonomy

RankClassification
Common NameGreenland Shark
DomainEukarya
KingdomAnimalia
PhylumChordata
ClassChondrichthyes
OrderSqualiformes
FamilySomniosidae
GenusSomniosus
SpeciesSomniosus microcephalus

2. Habitat

Distribution map of the Greenland shark

Where Greenland sharks are found

Greenland sharks live in the cold North Atlantic and Arctic Oceans, around Greenland, Iceland, Canada, and Norway. They've also been spotted as far south as the Gulf of Mexico following deep cold currents (Edwards et al., 2022). They usually live between 100 and 1,200 meters deep and prefer water between -1.1°C and 7.4°C, in summer they go deeper to stay cold and in winter they come closer to the surface (Watanabe et al., 2012).

3. Adaptive Traits

Close-up of a Greenland shark

Greenland shark up close

Trait 1: Extreme Longevity

Greenland sharks are the longest-living vertebrates with estimated lifespans of 272 to 512 years. They don't reach sexual maturity until around 150 years old. The freezing water slows down their metabolism so much that they age extremely slowly and they can go long periods without eating (Nielsen et al., 2016). They swim at only 0.34 m/s on average which makes them the slowest fish on record for their size (Watanabe et al., 2012).

Trait 2: TMAO and Urea as Biological Antifreeze

Their flesh contains urea and trimethylamine N-oxide (TMAO) which act like antifreeze, keeping their cells and proteins stable in freezing water and under extreme pressure (MacNeil et al., 2012). This makes their meat toxic to eat raw, in Iceland the dish hakarl involves fermenting the meat for months to make it safe.

Trait 3: Hunting While Basically Blind

Most adults have a parasite called Ommatokoita elongata attached to their corneas which destroys their vision (Benz et al., 2002). They compensate with a strong sense of smell, a lateral line that detects pressure changes and ampullae of Lorenzini that sense electrical fields from other animals.

4. Reproductive Information

Mating Rituals

Greenland shark mating has never been directly observed due to their deep dark habitat. Females likely release pheromones to attract males from a distance and males use claspers which are modified pelvic fins for fertilization (Noell et al., 2020).

Live Birth or Eggs?

Greenland sharks have live birth. They're ovoviviparous so babies develop inside the mother using yolk for nutrients. Gestation is estimated at 8 to 18 years, litter sizes range from about 10 to potentially over 200 and pups are about 38-42 cm at birth (Nielsen et al., 2016; Noell et al., 2020).

5. Anatomical Structures

Greenland shark in deep water

Greenland shark in the deep ocean

Skeletal System

As a cartilaginous fish their skeleton is made of cartilage not bone. They have a stocky body with a short snout, tiny eyes and small fins. Their upper 48-52 teeth are thin and pointy for gripping while the lower 48-52 are wide and flat for cutting (Yano et al., 2004; Compagno, 1984).

Digestive System

They have a stretchy J-shaped stomach for large meals and a spiral valve intestine that increases surface area for absorbing nutrients. Their large liver stores squalene oil for buoyancy and to eat large prey they bite down and roll their body to rip off chunks (Leclerc et al., 2012; Compagno, 1984).

Respiratory System

They breathe through five pairs of gill slits. They also have spiracles behind their eyes that let them take in water without swimming, which helps since they spend a lot of time barely moving (MacNeil et al., 2012).

Circulatory System

They have a two-chambered heart. Blood flows from the heart to the gills for oxygen then through the body and back. Their hemoglobin doesn't bind oxygen as strongly as warm-water sharks but cold water holds more dissolved oxygen so it works out (MacNeil et al., 2012).

6. Social Patterns

Greenland sharks are solitary. Multiple sharks may gather at a food source like a whale carcass but they don't truly socialize, their movements are driven by water temperature and if they communicate at all it's likely through chemical signals (MacNeil et al., 2012; Edwards et al., 2022).

7. Activity Patterns

They cruise at 0.34 m/s and max out at 0.74 m/s which makes them the slowest fish on record for their size (Watanabe et al., 2012). They live in deep water where it's always dark so day and night don't really matter much, they move to shallower water in winter and deeper water in summer (Ste-Marie et al., 2022).

8. Feeding

What Does It Eat?

They eat fish like herring, salmon, cod and halibut, as well as marine mammals like seals and belugas and also squid. Some stomachs have even contained moose and reindeer remains which were likely carcasses that fell into the water. Young ones eat mostly squid and switch to fish and seals as they grow (Fisk et al., 2002; Nielsen et al., 2019).

How Does It Eat?

They use suction feeding, expanding their mouth to create a vacuum that pulls prey in. For large prey, they bite down and roll their body to tear off pieces (Leclerc et al., 2012; Compagno, 1984).

How Does It Get Its Food?

They scavenge by following the scent of decaying flesh and actively hunt by ambushing resting prey. Seals sometimes sleep on the ocean floor and these sharks move slowly enough to approach undetected (Leclerc et al., 2012; Ste-Marie et al., 2022).

10. Works Cited

  1. Benz, G. W., Borucinska, J. D., Lowry, L. F., & Whiteley, H. E. (2002). Ocular lesions associated with attachment of the copepod Ommatokoita elongata to corneas of Pacific sleeper sharks. Journal of Parasitology, 88(3), 474-481. https://doi.org/10.1645/0022-3395(2002)088[0474:OLAAWA]2.0.CO;2
  2. Compagno, L. J. V. (1984). Sharks of the world: An annotated and illustrated catalogue of shark species known to date. Part 1. Food and Agriculture Organization of the United Nations.
  3. Edwards, J. E., Hiltz, E., Broell, F., Bushnell, P. G., Campana, S. E., Christiansen, J. S., Devine, B. M., Gallant, J. J., Hedges, K. J., MacNeil, M. A., McMeans, B. C., Rast, J. P., Hussey, N. E., & Fisk, A. T. (2022). Multi-year acoustic tracking reveals transient movements, recurring hotspots, and apparent seasonality in the coastal-offshore presence of Greenland sharks. Frontiers in Marine Science, 9, Article 902854. https://doi.org/10.3389/fmars.2022.902854
  4. Fisk, A. T., Tittlemier, S. A., Pranschke, J. L., & Norstrom, R. J. (2002). Using anthropogenic contaminants and stable isotopes to assess the feeding ecology of Greenland sharks. Ecology, 83(8), 2162-2172. https://doi.org/10.1890/0012-9658(2002)083[2162:UACASI]2.0.CO;2
  5. Hussey, N. E., MacNeil, M. A., McMeans, B. C., Olin, J. A., Dudley, S. F. J., Cliff, G., Wintner, S. P., Fennessy, S. T., & Fisk, A. T. (2014). Rescaling the trophic structure of marine food webs. Ecology Letters, 17(2), 239-250. https://doi.org/10.1111/ele.12226
  6. Leclerc, L. M., Lydersen, C., Haug, T., Bachmann, L., Fisk, A. T., & Kovacs, K. M. (2012). A missing piece in the Arctic food web puzzle? Inferred predation on belugas by Greenland sharks in Svalbard, Norway. Polar Biology, 35(8), 1197-1208. https://doi.org/10.1007/s00300-012-1166-7
  7. MacNeil, M. A., McMeans, B. C., Hussey, N. E., Vecsei, P., Svavarsson, J., Kovacs, K. M., Lydersen, C., Treble, M. A., Skomal, G. B., Ramsey, M., & Fisk, A. T. (2012). Biology of the Greenland shark Somniosus microcephalus. Journal of Fish Biology, 80(5), 991-1018. https://doi.org/10.1111/j.1095-8649.2012.03257.x
  8. Nielsen, J., Hedeholm, R. B., Heinemeier, J., Bushnell, P. G., Christiansen, J. S., Olsen, J., Ramsey, C. B., Brill, R. W., Simon, M., Steffensen, K. F., & Steffensen, J. F. (2016). Eye lens radiocarbon reveals centuries of longevity in the Greenland shark (Somniosus microcephalus). Science, 353(6300), 702-704. https://doi.org/10.1126/science.aaf1703
  9. Nielsen, J., Christiansen, J. S., Gronkjaer, P., Bushnell, P., Steffensen, J. F., Kiilerich, H. O., Praebel, K., & Hedeholm, R. (2019). Greenland shark (Somniosus microcephalus) stomach contents and stable isotope values reveal an ontogenetic dietary shift. Frontiers in Marine Science, 6, 125. https://doi.org/10.3389/fmars.2019.00125
  10. Noell, S., Hussey, N. E., & McMeans, B. C. (2020). Viviparity in the longest-living vertebrate, the Greenland shark (Somniosus microcephalus). Polar Biology, 43, 1459-1465. https://doi.org/10.1007/s00300-020-02727-3
  11. Ste-Marie, E., Watanabe, Y. Y., Semmens, J. M., Heard, M., & Hussey, N. E. (2022). Life in the slow lane: field metabolic rate and prey consumption rate of the Greenland shark modelled using archival biologgers. Journal of Experimental Biology, 225(7), jeb242994. https://doi.org/10.1242/jeb.242994
  12. Watanabe, Y. Y., Lydersen, C., Fisk, A. T., & Kovacs, K. M. (2012). The slowest fish: Swim speed and tail-beat frequency of Greenland sharks. Journal of Experimental Marine Biology and Ecology, 426-427, 5-11. https://doi.org/10.1016/j.jembe.2012.05.021
  13. Yano, K., Stevens, J. D., & Compagno, L. J. V. (2004). A review of the systematics of the sleeper shark genus Somniosus. Ichthyological Research, 51(4), 360-373. https://doi.org/10.1007/s10228-004-0244-4